'Queen' Gramma the Tortoise, the Oldest Animal at the San Diego Zoo, Dies at the Estimated Age of 141

Gramma the Galápagos tortoise died on Nov. 20, 2025. The oldest animal at the San Diego Zoo was believed to be around 141 ...

Gramma the Galapagos tortoise, oldest resident of San Diego Zoo, dies at about 141

The beloved Galápagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has ...

Gramma, San Diego Zoo’s ‘queen’ tortoise, dies at an estimated age of 141

Gramma, a Galápagos tortoise and the oldest resident of the San Diego Zoo, died Nov. 20. Her estimated 141-year lifespan spanned two world wars and 20 U.S. presidents.

Gramma, San Diego Zoo’s Galápagos tortoise, dies at 141

She had “ongoing bone conditions related to advanced age” that recently progressed and the care team “made the compassionate ...
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Gramma, the historic Galapagos tortoise at the San Diego Zoo who lived for more than a century, has died

Gramma, the San Diego Zoo's most iconic Galapagos tortoise, died at the age of 141 following a compassionate decision by the ...
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San Diego Zoo's oldest resident, Galápagos tortoise 'Gramma' dies

San Diego Zoo staff and animal lovers are reeling this week following the death of beloved Gramma the Galápagos tortoise, by ...
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141-Year-Old Galápagos Tortoise Gramma Dies at San Diego Zoo

A 141-year-old tortoise, celebrated as a star attraction at the San Diego Zoo in the United States and born in the 19th ...