'Queen' Gramma the Tortoise, the Oldest Animal at the San Diego Zoo, Dies at the Estimated Age of 141

Gramma the Galápagos tortoise died on Nov. 20, 2025. The oldest animal at the San Diego Zoo was believed to be around 141 ...

Gramma the Galapagos tortoise, oldest resident of San Diego Zoo who delighted visitors for almost a century, dies at about 141

LOS ANGELES (AP) — After more than a century of munching on her favorite foods of romaine lettuce and cactus fruit, beloved ...

Oldest-Resident Galapagos Tortoise at San Diego Zoo Dies at 141 and Had the Most Fitting Name

Speaking of long lives, the San Diego Zoo is saying goodbye to their oldest living resident, a Galapagos tortoise named ...
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San Diego Zoo's oldest resident, Galápagos tortoise 'Gramma' dies

San Diego Zoo staff and animal lovers are reeling this week following the death of beloved Gramma the Galápagos tortoise, by ...
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141-Year-Old Galápagos Tortoise Gramma Dies at San Diego Zoo

A 141-year-old tortoise, celebrated as a star attraction at the San Diego Zoo in the United States and born in the 19th ...

Gramma, San Diego Zoo’s Galápagos tortoise, dies at 141

She had “ongoing bone conditions related to advanced age” that recently progressed and the care team “made the compassionate ...